I'm currently trying out Joplin after having been using the Zim Desktop Wiki for several years. One minor feature I'm missing from Zim is that there were keyboard shortcuts for heading formatting. CTRL-1, CTRL-2, etc. would apply a heading level from 1 to the maximum supported heading level.
Also, a minor note: the wysiwyg editor has buttons for up to three heading levels, but markdown/joplin supports up to four levels.